The 25-year-old hip-hop superstar was shot dead in Las Vegas in 1996.
Duane "Keffe D" Davis, a former gang leader, told detectives in 2008 that his nephew Orlando "Baby Lane" Anderson had fired the shots.
At the time of the interview, his nephew had been dead for a decade.
Davis said the white Cadillac he and Anderson were riding in with two other men pulled up alongside a car with Shakur inside, along with hip-hop mogul and Death Row Records head Marion "Suge" Knight.
"If we would have been on my side, I would have blasted," Davis said. Instead, Knight's car was on the other side, so Davis passed his gun back to Deandre "Big Dre" Smith. But Smith didn't want it, Davis said.
Anderson, who was also in the back seat, took the weapon and started firing, Davis said.
Jurors watched the screen as the transcript of the audio recording rolled.
The interview is essential evidence in the state's case against the 63-year-old Davis. Prosecutors told jurors in opening statements that they would lean heavily on Davis' own admissions in interviews with police and on podcasts, as well as the 2019 memoir he co-authored, "Compton Street Legend."
He also previously told the FBI that his nephew was not involved in the 1996 shooting of Shakur, and speculated that Knight or Sean "Diddy" Combs were involved.
Davis has pleaded not guilty. His lawyer has called the prosecution's narrative "fiction" and said Davis was "full of crap" when he told stories of his involvement, and that authorities knew it. He argued investigators failed to back up the statements his client made with evidence.
The Los Angeles police detectives had actually brought Davis in to talk about the 1997 shooting death of Shakur's archrival, the Notorious B.I.G.
Davis had briefly been one of the suspects in that still unsolved killing, but he was later discounted.
Prosecutors allege Shakur was killed to avenge Shakur's entourage attacking Davis' nephew Anderson a few hours earlier at the MGM Grand after a Mike Tyson fight. Davis was at the fight but not the brawl.
Later in the evening, they were in the Cadillac and saw Shakur hanging out of a black BMW with people calling out "Tupac!"
Davis said that after Lane fired the shots, he thought Knight was dead.
Decades after his death, Shakur remains a cultural icon, and one of the most influential and versatile rappers of all time. He died as his popularity was peaking and his film career as an actor was taking off. His legend has only grown in the years since.
12°C